|1984||M.S. in Biochemistry (Honor Diploma), Moscow State University, Moscow, Russia|
|1984-1990||Ph.D. in Molecular Biology, Moscow State University, Moscow, Russia.
Thesis Advisor: Professor Alexei A. Bogdanov.
Postgraduate Training and Fellowship Appointments:
|1991 – 1995||Postdoctoral Fellow, Department of Biochemistry, Roche Institute of Molecular Biology, Nutley, NJ. Supervisor: Dr. James E. Ofengand.|
|1995 – 1999||Postdoctoral Fellow, Department of Developmental Neurobiology, St. Jude Children's Research Hospital, Memphis, TN 38105. Supervisor: Dr. Tom Curran.|
|1999 – 2003||Research Assistant Professor, Department of Medicine, Vanderbilt University School of Medicine, Nashville, TN; Director of the research program: Dr. Carlos Arteaga|